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一是通過過谷氨酸途徑,二是通過鳥氨酸途徑。其中,谷氨酸途徑在滲透脅迫和氮素不足情況下占主要地位;鳥氨酸途徑在氮素充足情況下占主導地位。谷氨酸途徑中,谷氨酸(Glu)在吡咯啉-5-羧酸合成酶(P5CS)催化下生成谷氨酰半醛(GSA),GSA自動環化為吡咯啉-5-羧酸(P5C),然后P5C在吡咯啉5-羧酸還原酶(P5CR)作用下生成脯氨酸。而鳥氨酸途徑與谷氨酸途徑在反應上的不同僅僅是第一步反應的底物和酶發生了變化,即由Glu和P5CS換成了鳥氨酸(Orn)和鳥氨酸轉氨酶(OAT)。脯氨酸的降解過程基本上是合成途徑的逆轉。脯氨酸首先被脯氨酸脫氫酶(PDH)氧化成P5C,后者在吡咯啉-5-羧酸脫氫酶的作用下生成Glu。高等植物體內脯氨酸合成與降解途徑.

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One through the glutamate pathway and the other through the ornithine pathway. Glutamate pathway plays the main role in osmotic stress and nitrogen deficiency. The ornithine pathway plays a dominant role in nitrogen abundance. In the glutamate pathway, glutamic acid (Glu) is catalyzed by pyrropridine 5-carboxylic acid synthetase (P5CS) to generate glutamyl semi-aldehyde (GSA), which is automatically cycled to pyrropridine 5-carboxylic acid (P5C), and then P5C is cycled to proline under the action of pyrropridine 5-carboxylic acid reductase (P5CR). The difference between the ornithine pathway and the glutamate pathway in the reaction is only that the substrate and enzyme of the first step of the reaction change, namely, from Glu and P5CS to ornithine (Orn) and ornithine transaminase (OAT). The degradation of proline is basically a reversal of the synthetic pathway. Proline is first oxidized to P5C by proline dehydrogenase (PDH), which generates Glu under the action of pyrrolidine -5- carboxylic acid dehydrogenase. Pathway of proline synthesis and degradation in higher plants.


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